Thank you, everyone.
Finally, I found a solution.
Step 1:
app.UseTwitterAuthentication(new TwitterAuthenticationOptions()
{
ConsumerKey = "XXX",
ConsumerSecret = "XXX",
BackchannelCertificateValidator = new Microsoft.Owin.Security.CertificateSubjectKeyIdentifierValidator(new[]
{
"A5EF0B11CEC04103A34A659048B21CE0572D7D47", // VeriSign Class 3 Secure Server CA - G2
"0D445C165344C1827E1D20AB25F40163D8BE79A5", // VeriSign Class 3 Secure Server CA - G3
"7FD365A7C2DDECBBF03009F34339FA02AF333133", // VeriSign Class 3 Public Primary Certification Authority - G5
"39A55D933676616E73A761DFA16A7E59CDE66FAD", // Symantec Class 3 Secure Server CA - G4
"add53f6680fe66e383cbac3e60922e3b4c412bed", // Symantec Class 3 EV SSL CA - G3
"4eb6d578499b1ccf5f581ead56be3d9b6744a5e5", // VeriSign Class 3 Primary CA - G5
"5168FF90AF0207753CCCD9656462A212B859723B", // DigiCert SHA2 High Assurance Server CA
"B13EC36903F8BF4701D498261A0802EF63642BC3" // DigiCert High Assurance EV Root CA
}),
});
Run the project again. I met the error as below:

Step 2:
https://apps.twitter.com/
Go to the Settings Tab Set a Callback URL
to any website. Even if it is not real
=> I set it to http://google.com

Then, It works for me. I don't know why I need to set Callback URL. After I set any value into it, I cannot set empty value. So, I think it's an error of Twitter API.
asp-net-mvc-5-owin-twitter-auth-throwing-401-exception