I am having same error, but my case is different. Please let me know different things below.
The email id for the request certificate should be same as the email id of iOS developer account and should be same as iTunes uploading account.
in my case the id which i bind with request certificate (abc@abc.com) is same as dev account (login to dev site for creating profile and all - abc@abc.com) and its different when i upload to iTunes (xyz@abc.com)
Please help me in this case, here are some confusion, and due to this I am getting this above subject error.
There is no invalid code sign it comes when I upload to iTunes via app loader.